Class Schedules

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q2

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:
Writing WC
Quantitative Thinking Q2

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q2

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

How do we observe, interpret, and understand the diverse world of living things that we see all around us? Students in our collaborative First-Year Course will explore this question from historical, artistic, and scientific perspectives. The Enlightenment approach to naming, categorizing, and representing the diversity of life dominates current academic science. Military activity enabled the work of key founding figures of this tradition; e.g., Charles Darwin’s critical insights were made as part of a British naval expedition whose ultimate goal was to maintain and expand colonial power. This approach to understanding nature has been practiced through striking traditions of drawing, printmaking, taxidermy, and cabinets of curiosities. Students will confront the practices and institutions of the biodiversity sciences in historical and political context. We will engage with indigenous counter-narratives and approaches to understanding nature outside of normative European colonial practices. Finally, we will work together to think critically about how we might decolonize institutions such as museums of natural history, approaches to visual culture and the biodiversity sciences more generally. Course activities will include active visual interpretation of nature through the production of artistic works, fieldtrips to museums such as the Minnesota Insect Collection and the Bell Museum of Natural History, and interpretation of archival materials such as Macalester’s Encyclopedie (Diderot), the Wangensteen Library’s Surinaamsche insecten engravings by Maria Sibylla Merian or the Science Museum of Minnesota’s Amazonian collection. One possible culminating project would be for students to develop a decolonial cabinet of curiosities in collaboration with the Macalester archives.

General Education Requirements:
Writing WA

Distribution Requirements:
Natural science and mathematics

Course Materials

Cells are the foundational organizing units of all life on our planet.  In this course, we study the ways in which these nanomachines use the shapes and interactions of molecules to store and convert information, energy, and mass, and to communicate with their environments.  The workings of single-celled bacteria to multicellular organisms, and the complex behaviors of multi-species ecosystems all depend on the working of these miniscule molecular gears packed inside cells. This course covers the structures and behavior of biological molecules, the metabolic reactions that power life, the properties of the fluid, complex membranes that organize biological molecules in space and time, the ways in which information is sent and received within and between cells, and molecular-scale understandings of health and disease processes.  Three lecture hours each week. Prerequisite(s): BIOL 190; and CHEM 112 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the science of invertebrate zoology. The vast majority of animals are invertebrates, including beautiful and charismatic organisms such as corals and butterflies, and also pests and parasites such as mosquitoes and tapeworms. Students will become familiar with all major and some minor phyla of marine, terrestrial, and freshwater animals. Through lectures, discussions, field trips, dissections, and laboratory observations of live organisms students will learn to identify invertebrates and understand their anatomy, life cycles, and evolutionary history. Students will complete independent projects involving field collection and identification of either insects or shells (mollusks). Three lecture hours and one three-hour lab each week. Prerequisite(s): BIOL 180

General Education Requirements:
Writing WA

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the science of invertebrate zoology. The vast majority of animals are invertebrates, including beautiful and charismatic organisms such as corals and butterflies, and also pests and parasites such as mosquitoes and tapeworms. Students will become familiar with all major and some minor phyla of marine, terrestrial, and freshwater animals. Through lectures, discussions, field trips, dissections, and laboratory observations of live organisms students will learn to identify invertebrates and understand their anatomy, life cycles, and evolutionary history. Students will complete independent projects involving field collection and identification of either insects or shells (mollusks). Three lecture hours and one three-hour lab each week. Prerequisite(s): BIOL 180

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

The structure and function of the vertebrate nervous system, with a focus on the human nervous system, will be explored through analysis of human brain specimens and sections, and artistic photographic and computer graphic representations of nervous system structures at both the microscopic and systems levels. This course is designed to provide an understanding of both the peripheral and central nervous system with a focus on brain, brainstem, and spinal cord structure and function through observation and study of normal brain tissue and discussion of clinical cases. This course is intended for students with a strong interest in neuroscience. Three lecture hours and one three-hour lab each week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

The structure and function of the vertebrate nervous system, with a focus on the human nervous system, will be explored through analysis of human brain specimens and sections, and artistic photographic and computer graphic representations of nervous system structures at both the microscopic and systems levels. This course is designed to provide an understanding of both the peripheral and central nervous system with a focus on brain, brainstem, and spinal cord structure and function through observation and study of normal brain tissue and discussion of clinical cases. This course is intended for students with a strong interest in neuroscience. Three lecture hours and one three-hour lab each week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

As a field, agroecology considers agricultural landscapes in the context of ecological principles and concepts. We will investigate the ecological underpinnings of agriculture, including interactions between soils, microbes, plants and animals, always in the context of climate change, land use change and other global change drivers. In addition to exploring the water and nutrient demands of agricultural systems from a physiological perspective and conventional agricultural systems, we will also discuss sustainable agricultural practices and sustainability in the global food system. This class will feature case studies from around the globe. Prerequisite(s): BIOL 170 or permission of instructor.

General Education Requirements:

Distribution Requirements:

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

The principles of genetics explain how the information in the genome is used to build cells and organisms, how traits are inherited, and how mutations can lead to genetic diseases and cancer but also evolutionary innovation and new species. This course provides an in-depth study of the principles of modern genetics and will cover gene mapping of Mendelian and non-Mendelian traits as well as concepts and methodologies of molecular genetics, including gene expression analysis, next generation sequencing, and gene editing. We will also explore the field of epigenetics and discuss the mechanisms by which nature and nurture together affect the phenotype of an individual. Finally, we will discuss bioethical and sociocultural issues related to the application of genetic principles such as genetic testing, genome manipulation, and gene therapy. Three lecture hours each week. Prerequisite(s): BIOL 190

General Education Requirements:
Quantitative Thinking Q1

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course is an introduction to vertebrate immunity. Its evolution, cellular and molecular mechanisms, health and disease functions and therapeutic manipulations are explored through approaches including lectures, clinical case studies, extensive reading of the primary literature, problem-solving and an intensive focus on scientific writing. The course typically includes a civic engagement component where students work with health organizations and schools in the Twin Cities area. The course laboratory uses guided exercises and independent projects to focus on current immunological techniques including flow cytometry, magnetic cell sorting and antibody assays. Three hours of lectures and four to six hours of laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200; CHEM 112; junior or senior standing

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course is an introduction to vertebrate immunity. Its evolution, cellular and molecular mechanisms, health and disease functions and therapeutic manipulations are explored through approaches including lectures, clinical case studies, extensive reading of the primary literature, problem-solving and an intensive focus on scientific writing. The course typically includes a civic engagement component where students work with health organizations and schools in the Twin Cities area. The course laboratory uses guided exercises and independent projects to focus on current immunological techniques including flow cytometry, magnetic cell sorting and antibody assays. Three hours of lectures and four to six hours of laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200; CHEM 112; junior or senior standing

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Ecology and environmental science are increasingly using 'big data' to expand and refine research questions. We will examine, analyze, and interpret datasets that represent a wide range of ecological topics and approaches, including nutrient cycling, hydrology, climate change, human and animal migration, satellite remote sensing, and biodiversity. The course will examine recent literature and apply novel analyses using open-access data and code every week. We will build skills in R programming, science communication, data visualization, and critical examination of literature. The course is project-oriented and students will work independently and in small groups to dive deeply into large data using R/RStudio, and produce original analyses and results. Three lecture hours and three hours of laboratory each week. Prerequisite(s): BIOL 170; and STAT 112 or STAT 155

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Ecology and environmental science are increasingly using 'big data' to expand and refine research questions. We will examine, analyze, and interpret datasets that represent a wide range of ecological topics and approaches, including nutrient cycling, hydrology, climate change, human and animal migration, satellite remote sensing, and biodiversity. The course will examine recent literature and apply novel analyses using open-access data and code every week. We will build skills in R programming, science communication, data visualization, and critical examination of literature. The course is project-oriented and students will work independently and in small groups to dive deeply into large data using R/RStudio, and produce original analyses and results. Three lecture hours and three hours of laboratory each week. Prerequisite(s): BIOL 170; and STAT 112 or STAT 155

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course aims to integrate organismal, cellular, genetic and molecular approaches to the study of animal development. We will analyze a diversity of mechanisms, ranging from ones that set up pattern formation in the unfertilized egg to those governing morphogenesis of organ systems. Evolution of developmental mechanisms will also be discussed. The lab component will incorporate both descriptive and experimental embryological techniques. Three hours of lecture and one three-hour laboratory each week. Prerequisite(s): BIOL 180, BIOL 190 and BIOL 200, or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course aims to integrate organismal, cellular, genetic and molecular approaches to the study of animal development. We will analyze a diversity of mechanisms, ranging from ones that set up pattern formation in the unfertilized egg to those governing morphogenesis of organ systems. Evolution of developmental mechanisms will also be discussed. The lab component will incorporate both descriptive and experimental embryological techniques. Three hours of lecture and one three-hour laboratory each week. Prerequisite(s): BIOL 180, BIOL 190 and BIOL 200, or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Why do animals behave the way they do? Why do lions have manes while leopards don’t? Why do elephants and bees live in groups but many other species do not? Why does your friendly neighborhood squirrel get so busy late in the fall and again in the spring? Why do certain wolves ‘fish’ but others never learn the technique? In this course we will explore the fundamentals of animal behavior and use that foundation to understand how we can better manage and conserve animal biodiversity. Labs will include hands-on experience with tools that will allow us to quantify behavior, develop ethograms, and understand species’ repertoires. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisites: BIOL/ENVI 170 and either BIOL180 or ENVI 240. Recommended: STAT 155.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Why do animals behave the way they do? Why do lions have manes while leopards don’t? Why do elephants and bees live in groups but many other species do not? Why does your friendly neighborhood squirrel get so busy late in the fall and again in the spring? Why do certain wolves ‘fish’ but others never learn the technique? In this course we will explore the fundamentals of animal behavior and use that foundation to understand how we can better manage and conserve animal biodiversity. Labs will include hands-on experience with tools that will allow us to quantify behavior, develop ethograms, and understand species’ repertoires. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisites: BIOL/ENVI 170 and either BIOL180 or ENVI 240. Recommended: STAT 155.

General Education Requirements:

Distribution Requirements:

Course Materials

This course examines neural control of movement through the lens of the formative primary literature with an emphasis on computational techniques. Students will integrate basic science and neurophysiology with quantitative approaches to understand electrophysiology of the motor cortex, basal ganglia, and cerebellum, as well as translational work such as neural prosthetics and deep brain stimulation. Prerequisite: BIOL 200 and PSYC 248.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

As we usher into the Anthropocene, where ‘Amazon’ is recognized as a global marketing forum more than the mighty river, the fate of biodiversity reflects a similar trajectory. Human population growth has resulted in unprecedented changes in our planet’s ecosystems. Species have been lost, modified, and forced to live in concrete jungles. Human history, culture and socio-political nuances in different parts of the world result in distinctive challenges as well as enigmatically optimistic scenarios with respect to wildlife conservation. In this seminar we will investigate perspectives from South Asia where animals and humans coexist at very high density and proximity, consider the North American system where views about biodiversity can often be very polarizing, and explore conservation in Africa where pockets of pristine wilderness persist. We will engage with the primary literature of wildlife conservation, hear from scientists on the frontlines, and discuss the idea that there could be a unifying model that safeguards biodiversity without detrimentally impeding human development. Three hours of lecture/discussion each week. Prerequisites: BIOL/ENVI 170 and either BIOL 180 or ENVI 240.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

What exactly happens inside the brain when the mind goes awry? What are the subjective experiences of psychiatric disorders and how can our understanding of the brain better elucidate the causes and potential treatments for these conditions? Today, modern neuroscience has significantly advanced our insight into these questions. This course is designed to overview major types of mental illness and delve into their origins using a neuroscience approach. In order to explore these topics, class will involve (1) first-hand narratives of individuals with mental illness, (2) interactive lectures, during which you’ll acquire and share knowledge, (3) group presentations, and (4) group discussions on neuroscience research. As students in this course, you will be evaluated through written and presented work, as well as critical review and discussion of neuroscience research. By the completion of this course, you should have a detailed understanding of the neurobiology of psychopathologies, and an increased capacity to read and interpret neuroscience research. Prerequisite: PSYC 201 or STAT 155; and PSYC 244 or PSYC 246 or PSYC 248, or BIOL 316 with PSYC 100 or PSYC 180.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course will deal with aspects of human anatomy and physiology of special interest to women and/or those who identify as women, especially relating to sexuality and reproduction. Biological topics covered will include menstruation and menopause, sexuality, conception, contraception, infertility, abortion, pregnancy, cancer, and AIDS. Advances in assisted reproductive technologies, hormone therapies, and genetic engineering technologies will be discussed. Three lecture hours each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q3

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:
Quantitative Thinking Q2

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q1

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Cells are the foundational organizing units of all life on our planet.  In this course, we study the ways in which these nanomachines use the shapes and interactions of molecules to store and convert information, energy, and mass, and to communicate with their environments.  The workings of single-celled bacteria to multicellular organisms, and the complex behaviors of multi-species ecosystems all depend on the working of these miniscule molecular gears packed inside cells. This course covers the structures and behavior of biological molecules, the metabolic reactions that power life, the properties of the fluid, complex membranes that organize biological molecules in space and time, the ways in which information is sent and received within and between cells, and molecular-scale understandings of health and disease processes.  Three lecture hours each week. Prerequisite(s): BIOL 190; and CHEM 112 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A continuation of BIOL 351. A study of biological processes at the molecular level with an emphasis on the metabolism of amino acids, nucleotides, the regulation of biochemical pathways, and topics in molecular biology such as gene replication, the synthesis of proteins and nucleic acids, and recent advances in genomics and proteomics. Three lecture hours each week. Prerequisite(s): BIOL 190, BIOL 200, and BIOL 351, or permission of instructor.

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Distribution Requirements:
Natural science and mathematics

Course Materials

How are therapeutic and diagnostic agents designed? How are these chemicals used to detect and treat illnesses? In this class, we aim to answer these questions by exploring molecular medicine from a (bio)chemical perspective. We will discuss the chemical basis of different diseases and how they can be targeted by therapeutics and monitored using chemical diagnostics. We will examine the fundamentals of drug design and development and dive into primary literature to explore cutting edge research in the field of medicinal chemistry. Prerequisite(s): BIOL 351 (or concurrent registration)

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Vertebrates are backboned organisms that include more than 50,000 living species ranging in size from a fish weighing less than an aspirin to a whale weighing nearly 300,000 pounds. They live in virtually every habitat on Earth and include species that fly higher, swim deeper, and move faster than other organisms. In this course, we will investigate vertebrate form and function through the lens of evolutionary history and dissection of representative vertebrates. Emphasis is placed on the origin and diversification of the basic vertebrate body plan and the morphological, functional, and evolutionary patterns that result. This course fulfills the coursework in anatomy recommended for most pre-health programs. Three lecture hours and three hours of dissection-based laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior standing or permission of instructor

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Vertebrates are backboned organisms that include more than 50,000 living species ranging in size from a fish weighing less than an aspirin to a whale weighing nearly 300,000 pounds. They live in virtually every habitat on Earth and include species that fly higher, swim deeper, and move faster than other organisms. In this course, we will investigate vertebrate form and function through the lens of evolutionary history and dissection of representative vertebrates. Emphasis is placed on the origin and diversification of the basic vertebrate body plan and the morphological, functional, and evolutionary patterns that result. This course fulfills the coursework in anatomy recommended for most pre-health programs. Three lecture hours and three hours of dissection-based laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior standing or permission of instructor

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

How are ecosystem carbon stocks responding to climate change? What controls primary production? How is agricultural land use change altering the nitrogen cycle? How do ecosystems respond to elevated atmospheric CO2 concentrations, and how does nutrient availability affect this CO2 response? What is the relationship between biodiversity and ecosystem function? These are all questions that ecosystem ecologists seek to answer. In this course, we will investigate the principles and processes that govern the structure and function of ecosystems, with an emphasis on how nutrients, water and energy cycle through ecosystems. Ecosystem ecology is interdisciplinary in nature, and draws from fields such as physiological, microbial and community ecology, soil science, atmospheric science, and geology. We will cover both fundamental principles and recent, cutting-edge research that focuses on global change drivers (e.g., climate change, nitrogen deposition, land use change, and altered disturbance regimes). Includes 3 hours of lab per week. Prerequisite(s): STAT 155 and BIOL 170. Recommended: CHEM 111 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

How are ecosystem carbon stocks responding to climate change? What controls primary production? How is agricultural land use change altering the nitrogen cycle? How do ecosystems respond to elevated atmospheric CO2 concentrations, and how does nutrient availability affect this CO2 response? What is the relationship between biodiversity and ecosystem function? These are all questions that ecosystem ecologists seek to answer. In this course, we will investigate the principles and processes that govern the structure and function of ecosystems, with an emphasis on how nutrients, water and energy cycle through ecosystems. Ecosystem ecology is interdisciplinary in nature, and draws from fields such as physiological, microbial and community ecology, soil science, atmospheric science, and geology. We will cover both fundamental principles and recent, cutting-edge research that focuses on global change drivers (e.g., climate change, nitrogen deposition, land use change, and altered disturbance regimes). Includes 3 hours of lab per week. Prerequisite(s): STAT 155 and BIOL 170. Recommended: CHEM 111 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course will cover research design and data analysis across biological sub-disciplines, with an emphasis on practical application. Research design topics will include basic and specialized experimental designs and avoiding pseudoreplication. Data analysis topics will focus primarily on statistical models: linear and non-linear, variable selection, and handling broken assumptions. Special topics may include multivariate methods, machine learning, and information theoretic approaches. The major aim of the course is to have students gain competence and confidence in the analysis of biological data - selecting appropriate statistical methods for particular questions and datasets, avoiding pitfalls, and contributing usefully and transparently to scientific knowledge. Three lecture hours per week. Prerequisites: STAT 155 and BIOL 170, 180, 190, or 200.

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Course Materials

Biodiversity is fast disappearing from the face of the planet. To keep a check on wildlife species, populations, and individuals, we need robust methods to enumerate individual and population scale processes such as abundance, distribution, resource use, and behavior. In this course you will be exposed to the fundamentals of wildlife monitoring techniques and their relevant applications. We will learn about wildlife census techniques and how these methods have expanded and championed wildlife conservation. We will engage in hands-on, outdoor experiences such as camera trapping, distance sampling, diet monitoring, animal capture and telemetry. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisites: BIOL/ENVI 170 and either BIOL 180 or ENVI 240; Junior or Senior Standing or permission of the instructor. Recommended: STAT 155.

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Course Materials

Varies by semester. Consult the department or class schedule for current listing.

General Education Requirements:

Distribution Requirements:

Course Materials

*ACTC students require permission of instructor

General Education Requirements:
Writing WA

Distribution Requirements:
Natural science and mathematics

Course Materials

Recent advances in molecular biology have lead to the ability to directly manipulate genomes in highly precise ways. CRISPR-based genome editing is a type of genetic engineering that can be used to add, replace, or delete specific sequences in an organism's genes using specialized nucleases.  The possible applications of this methodology are far-reaching and have already been used to correct disease-causing mutations in mouse models and human cells, leading to gene therapy clinical trials and the first reported case of manipulation of human embryos. We will discuss the primary literature from this emerging field, technical hurdles that may hamper medical applications of this technology such as "off-target" effects, and ethical and socio-cultural issues surrounding genome modification. Three lecture/discussion hours each week. Prerequisite(s): BIOL 190 and BIOL 200; junior or senior standing; or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

In this course students pursue a collaborative research project with a faculty member on a topic in contemporary biochemistry. In addition to extensive laboratory research, reading and discussion of the primary literature and oral and written presentation of research topics and results will be pursued. Three to six hours of lecture and/or laboratory each week. Depending on the semester/topic, this course may be taught be either Biology or Chemistry Faculty members. Prerequisite(s): BIOL 351; junior or senior standing; and permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course offers an opportunity to work with a faculty member on current research in animal diversity and evolution. Lab periods will be devoted to fieldwork, lab work, and data analysis. Lecture periods will focus on discussion of the scientific literature related to the research area undertaken. Research projects will be undertaken in the fields of population genetics, phylogenetic systematics, or biogeography. Techniques may include field collection of animals, species identification, DNA sequencing, analysis of genetic data sets, and scanning electron microscopy. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior or senior standing; and permission of instructor.

General Education Requirements:
Writing WA

Distribution Requirements:
Natural science and mathematics

Course Materials

This course offers an opportunity to work with a faculty member on current research in animal diversity and evolution. Lab periods will be devoted to fieldwork, lab work, and data analysis. Lecture periods will focus on discussion of the scientific literature related to the research area undertaken. Research projects will be undertaken in the fields of population genetics, phylogenetic systematics, or biogeography. Techniques may include field collection of animals, species identification, DNA sequencing, analysis of genetic data sets, and scanning electron microscopy. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior or senior standing; and permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This is an advanced course that examines current and emerging technologies for sensing and manipulation of neuronal populations in neuroscience research. Students will learn the scientific basis of techniques such as optogenetics and calcium imaging as well as their application via reading and discussing primary literature in systems neuroscience. Three lecture/discussion hours each week. Prerequisite(s): BIOL 200 and PSYC 248. (BIOL 316 recommended.)

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Disease is one of the major unpredictable factors that influences wildlife extinction. In the Anthropocene humans have altered habitats and temperature regimes, and have introduced novel organisms like domestic dogs and cats into natural systems. These changes predispose wildlife populations to become vulnerable to diseases while also facilitating the spill over of such diseases to humans. Deadly disease outbreaks such as the Bubonic Plague, Ebola, and even COVID-19 have not only caused human death and suffering but have crippled our economies and civilizations. In this seminar we will discuss major wildlife diseases, the ecology of their origin and repercussions on wildlife and humans. We will engage with the primary scientific literature, hear from scientists whose work is at the front lines of these spillovers, and learn from the mistakes that humans have made along the way. Three hours of lecture/discussion each week. Prerequisites: BIOL/ENVI 170 and either BIOL 180 or ENVI 240. Junior or Senior Standing or permission of the instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This seminar centers on our understanding of the molecular events underlying the development of various immunological disorders as well as other conditions, and the immunotherapeutic approaches for their treatment. Topics will cover autoimmunity, cancer immunotherapy, organ and stem cell transplantation, allergy, and vaccine development. Major goals of the course include developing an ability to understand immunological mechanisms behind these therapies on the molecular level and learning to critically evaluate existing literature in the field. Students will discuss primary articles and secondary reviews in a roundtable format, and will present written and oral critiques of the literature and develop a final individual project. Three lecture/discussion hours each week. Prerequisites: BIOL 200 (BIOL 357 recommended); junior or senior standing or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials