Class Schedules

This course will deal with aspects of human anatomy and physiology of special interest to women and/or those who identify as women, especially relating to sexuality and reproduction. Biological topics covered will include menstruation and menopause, sexuality, conception, contraception, infertility, abortion, pregnancy, cancer, and AIDS. Advances in assisted reproductive technologies, hormone therapies, and genetic engineering technologies will be discussed. Three lecture hours each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q3

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:
Quantitative Thinking Q2

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q1

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Cells are the foundational organizing units of all life on our planet.  In this course, we study the ways in which these nanomachines use the shapes and interactions of molecules to store and convert information, energy, and mass, and to communicate with their environments.  The workings of single-celled bacteria to multicellular organisms, and the complex behaviors of multi-species ecosystems all depend on the working of these miniscule molecular gears packed inside cells. This course covers the structures and behavior of biological molecules, the metabolic reactions that power life, the properties of the fluid, complex membranes that organize biological molecules in space and time, the ways in which information is sent and received within and between cells, and molecular-scale understandings of health and disease processes.  Three lecture hours each week. Prerequisite(s): BIOL 190; and CHEM 112 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A continuation of BIOL 351. A study of biological processes at the molecular level with an emphasis on the metabolism of amino acids, nucleotides, the regulation of biochemical pathways, and topics in molecular biology such as gene replication, the synthesis of proteins and nucleic acids, and recent advances in genomics and proteomics. Three lecture hours each week. Prerequisite(s): BIOL 190, BIOL 200, and BIOL 351, or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

How are therapeutic and diagnostic agents designed? How are these chemicals used to detect and treat illnesses? In this class, we aim to answer these questions by exploring molecular medicine from a (bio)chemical perspective. We will discuss the chemical basis of different diseases and how they can be targeted by therapeutics and monitored using chemical diagnostics. We will examine the fundamentals of drug design and development and dive into primary literature to explore cutting edge research in the field of medicinal chemistry. Prerequisite(s): BIOL 351 (or concurrent registration)

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Vertebrates are backboned organisms that include more than 50,000 living species ranging in size from a fish weighing less than an aspirin to a whale weighing nearly 300,000 pounds. They live in virtually every habitat on Earth and include species that fly higher, swim deeper, and move faster than other organisms. In this course, we will investigate vertebrate form and function through the lens of evolutionary history and dissection of representative vertebrates. Emphasis is placed on the origin and diversification of the basic vertebrate body plan and the morphological, functional, and evolutionary patterns that result. This course fulfills the coursework in anatomy recommended for most pre-health programs. Three lecture hours and three hours of dissection-based laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior standing or permission of instructor

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Vertebrates are backboned organisms that include more than 50,000 living species ranging in size from a fish weighing less than an aspirin to a whale weighing nearly 300,000 pounds. They live in virtually every habitat on Earth and include species that fly higher, swim deeper, and move faster than other organisms. In this course, we will investigate vertebrate form and function through the lens of evolutionary history and dissection of representative vertebrates. Emphasis is placed on the origin and diversification of the basic vertebrate body plan and the morphological, functional, and evolutionary patterns that result. This course fulfills the coursework in anatomy recommended for most pre-health programs. Three lecture hours and three hours of dissection-based laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior standing or permission of instructor

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

How are ecosystem carbon stocks responding to climate change? What controls primary production? How is agricultural land use change altering the nitrogen cycle? How do ecosystems respond to elevated atmospheric CO2 concentrations, and how does nutrient availability affect this CO2 response? What is the relationship between biodiversity and ecosystem function? These are all questions that ecosystem ecologists seek to answer. In this course, we will investigate the principles and processes that govern the structure and function of ecosystems, with an emphasis on how nutrients, water and energy cycle through ecosystems. Ecosystem ecology is interdisciplinary in nature, and draws from fields such as physiological, microbial and community ecology, soil science, atmospheric science, and geology. We will cover both fundamental principles and recent, cutting-edge research that focuses on global change drivers (e.g., climate change, nitrogen deposition, land use change, and altered disturbance regimes). Includes 3 hours of lab per week. Prerequisite(s): STAT 155 and BIOL 170. Recommended: CHEM 111 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

How are ecosystem carbon stocks responding to climate change? What controls primary production? How is agricultural land use change altering the nitrogen cycle? How do ecosystems respond to elevated atmospheric CO2 concentrations, and how does nutrient availability affect this CO2 response? What is the relationship between biodiversity and ecosystem function? These are all questions that ecosystem ecologists seek to answer. In this course, we will investigate the principles and processes that govern the structure and function of ecosystems, with an emphasis on how nutrients, water and energy cycle through ecosystems. Ecosystem ecology is interdisciplinary in nature, and draws from fields such as physiological, microbial and community ecology, soil science, atmospheric science, and geology. We will cover both fundamental principles and recent, cutting-edge research that focuses on global change drivers (e.g., climate change, nitrogen deposition, land use change, and altered disturbance regimes). Includes 3 hours of lab per week. Prerequisite(s): STAT 155 and BIOL 170. Recommended: CHEM 111 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course will cover research design and data analysis across biological sub-disciplines, with an emphasis on practical application. Research design topics will include basic and specialized experimental designs and avoiding pseudoreplication. Data analysis topics will focus primarily on statistical models: linear and non-linear, variable selection, and handling broken assumptions. Special topics may include multivariate methods, machine learning, and information theoretic approaches. The major aim of the course is to have students gain competence and confidence in the analysis of biological data - selecting appropriate statistical methods for particular questions and datasets, avoiding pitfalls, and contributing usefully and transparently to scientific knowledge. Three lecture hours per week. Prerequisites: STAT 155 and BIOL 170, 180, 190, or 200.

General Education Requirements:
Quantitative Thinking Q3

Distribution Requirements:

Course Materials

Biodiversity is fast disappearing from the face of the planet. To keep a check on wildlife species, populations, and individuals, we need robust methods to enumerate individual and population scale processes such as abundance, distribution, resource use, and behavior. In this course you will be exposed to the fundamentals of wildlife monitoring techniques and their relevant applications. We will learn about wildlife census techniques and how these methods have expanded and championed wildlife conservation. We will engage in hands-on, outdoor experiences such as camera trapping, distance sampling, diet monitoring, animal capture and telemetry. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisites: BIOL/ENVI 170 and either BIOL 180 or ENVI 240; Junior or Senior Standing or permission of the instructor. Recommended: STAT 155.

General Education Requirements:

Distribution Requirements:

Course Materials

Varies by semester. Consult the department or class schedule for current listing.

General Education Requirements:

Distribution Requirements:

Course Materials

*ACTC students require permission of instructor

General Education Requirements:
Writing WA

Distribution Requirements:
Natural science and mathematics

Course Materials

Recent advances in molecular biology have lead to the ability to directly manipulate genomes in highly precise ways. CRISPR-based genome editing is a type of genetic engineering that can be used to add, replace, or delete specific sequences in an organism's genes using specialized nucleases.  The possible applications of this methodology are far-reaching and have already been used to correct disease-causing mutations in mouse models and human cells, leading to gene therapy clinical trials and the first reported case of manipulation of human embryos. We will discuss the primary literature from this emerging field, technical hurdles that may hamper medical applications of this technology such as "off-target" effects, and ethical and socio-cultural issues surrounding genome modification. Three lecture/discussion hours each week. Prerequisite(s): BIOL 190 and BIOL 200; junior or senior standing; or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

In this course students pursue a collaborative research project with a faculty member on a topic in contemporary biochemistry. In addition to extensive laboratory research, reading and discussion of the primary literature and oral and written presentation of research topics and results will be pursued. Three to six hours of lecture and/or laboratory each week. Depending on the semester/topic, this course may be taught be either Biology or Chemistry Faculty members. Prerequisite(s): BIOL 351; junior or senior standing; and permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course offers an opportunity to work with a faculty member on current research in animal diversity and evolution. Lab periods will be devoted to fieldwork, lab work, and data analysis. Lecture periods will focus on discussion of the scientific literature related to the research area undertaken. Research projects will be undertaken in the fields of population genetics, phylogenetic systematics, or biogeography. Techniques may include field collection of animals, species identification, DNA sequencing, analysis of genetic data sets, and scanning electron microscopy. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior or senior standing; and permission of instructor.

General Education Requirements:
Writing WA

Distribution Requirements:
Natural science and mathematics

Course Materials

This course offers an opportunity to work with a faculty member on current research in animal diversity and evolution. Lab periods will be devoted to fieldwork, lab work, and data analysis. Lecture periods will focus on discussion of the scientific literature related to the research area undertaken. Research projects will be undertaken in the fields of population genetics, phylogenetic systematics, or biogeography. Techniques may include field collection of animals, species identification, DNA sequencing, analysis of genetic data sets, and scanning electron microscopy. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisite(s): BIOL 180 and BIOL 190; junior or senior standing; and permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This is an advanced course that examines current and emerging technologies for sensing and manipulation of neuronal populations in neuroscience research. Students will learn the scientific basis of techniques such as optogenetics and calcium imaging as well as their application via reading and discussing primary literature in systems neuroscience. Three lecture/discussion hours each week. Prerequisite(s): BIOL 200 and PSYC 248. (BIOL 316 recommended.)

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Disease is one of the major unpredictable factors that influences wildlife extinction. In the Anthropocene humans have altered habitats and temperature regimes, and have introduced novel organisms like domestic dogs and cats into natural systems. These changes predispose wildlife populations to become vulnerable to diseases while also facilitating the spill over of such diseases to humans. Deadly disease outbreaks such as the Bubonic Plague, Ebola, and even COVID-19 have not only caused human death and suffering but have crippled our economies and civilizations. In this seminar we will discuss major wildlife diseases, the ecology of their origin and repercussions on wildlife and humans. We will engage with the primary scientific literature, hear from scientists whose work is at the front lines of these spillovers, and learn from the mistakes that humans have made along the way. Three hours of lecture/discussion each week. Prerequisites: BIOL/ENVI 170 and either BIOL 180 or ENVI 240. Junior or Senior Standing or permission of the instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This seminar centers on our understanding of the molecular events underlying the development of various immunological disorders as well as other conditions, and the immunotherapeutic approaches for their treatment. Topics will cover autoimmunity, cancer immunotherapy, organ and stem cell transplantation, allergy, and vaccine development. Major goals of the course include developing an ability to understand immunological mechanisms behind these therapies on the molecular level and learning to critically evaluate existing literature in the field. Students will discuss primary articles and secondary reviews in a roundtable format, and will present written and oral critiques of the literature and develop a final individual project. Three lecture/discussion hours each week. Prerequisites: BIOL 200 (BIOL 357 recommended); junior or senior standing or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q3

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course dives into a range of topics to study how species, populations, communities, ecosystems, and biomes function in our changing climate. We will emphasize biological nutrient and energy cycling, population dynamics, animal and plant species interactions, disturbances and response to disturbances, and ecology in urban and agricultural landscapes. We will examine Ecology under four conceptual 'lenses': Climate Change, Environmental Justice, Land Use, and Ecosystem Services. These lenses provide critical insight into how scientists, policy makers, land managers, and other stakeholders evaluate complex ecological and environmental systems. Labs will be field and data-based, and emphasize the development of hypotheses, novel data collection at Ordway Field Station, and statistical analysis. Three hours lecture and one three-hour laboratory each week.

General Education Requirements:
Writing WA
Quantitative Thinking Q3

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the diversity and history of life. We begin with basic evolutionary patterns and processes that have shaped living things, and then move on to the major groups of organisms (their morphology, physiology, reproductive cycles) and their evolutionary origins and relationships. Using recent findings from such diverse fields as molecular phylogenetics, developmental biology, and paleontology, this course introduces students to the major branches on the tree of life. Three lecture hours and one three-hour lab each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:
Quantitative Thinking Q1

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An introduction to the principles of genetics, including patterns of inheritance, structure and function of genetic material, flow of genetic information, and control of gene expression. This course also introduces students to the methodologies used in genetics research, such as gene mapping, gene expression techniques, DNA sequencing, introductory genomics, and gene manipulation. Bioethical and sociocultural issues that emerge from the applications of genetic concepts and methodologies are discussed. Three lecture hours and one three-hour laboratory each week.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Cells are the foundational organizing units of all life on our planet.  In this course, we study the ways in which these nanomachines use the shapes and interactions of molecules to store and convert information, energy, and mass, and to communicate with their environments.  The workings of single-celled bacteria to multicellular organisms, and the complex behaviors of multi-species ecosystems all depend on the working of these miniscule molecular gears packed inside cells. This course covers the structures and behavior of biological molecules, the metabolic reactions that power life, the properties of the fluid, complex membranes that organize biological molecules in space and time, the ways in which information is sent and received within and between cells, and molecular-scale understandings of health and disease processes.  Three lecture hours each week. Prerequisite(s): BIOL 190; and CHEM 112 or CHEM 115

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course provides an introduction to outdoor education as an opportunity to promote social justice and environmental sustainability in a globalized world.  Informed by relevant philosophical, psychological, cultural and political-economic frameworks, in addition to critical issues in public education policy and practice, we will explore interdisciplinary approaches to outdoor environmental education appropriate for students across the K-12 continuum.  We will utilize the Katharine Ordway Natural History Study Area (Ordway Field Station) as an outdoor classroom and will adapt curriculum from the Minnesota Department of Natural Resources (DNR) and other outdoor education organizations to assist elementary school teachers and students in fulfilling Minnesota K-12 Academic Standards. Early in the semester, all students will participate in a weekend retreat at the Ordway Field Station. Weekly lab sessions will include field days during which course members design and implement educational experiences for elementary school children at Ordway, small group work days for preparing field day lesson plans, trips to local outdoor environmental education sites within the Twin Cities, and other experiential learning opportunities.  Weekly seminar sessions incorporating readings, reflective writing, and individual and small group projects complement the experiential aspects of the course. As the semester progresses, each course member will develop a curricular unit aimed at teaching an important environmental issue to diverse adolescents attending urban public schools.  The curricular unit is a significant undertaking that provides students with the opportunity to synthesize all aspects of the course material in a creative, pragmatic and integrative manner.

General Education Requirements:

Distribution Requirements:
Social science

Course Materials

This course provides an introduction to outdoor education as an opportunity to promote social justice and environmental sustainability in a globalized world.  Informed by relevant philosophical, psychological, cultural and political-economic frameworks, in addition to critical issues in public education policy and practice, we will explore interdisciplinary approaches to outdoor environmental education appropriate for students across the K-12 continuum.  We will utilize the Katharine Ordway Natural History Study Area (Ordway Field Station) as an outdoor classroom and will adapt curriculum from the Minnesota Department of Natural Resources (DNR) and other outdoor education organizations to assist elementary school teachers and students in fulfilling Minnesota K-12 Academic Standards. Early in the semester, all students will participate in a weekend retreat at the Ordway Field Station. Weekly lab sessions will include field days during which course members design and implement educational experiences for elementary school children at Ordway, small group work days for preparing field day lesson plans, trips to local outdoor environmental education sites within the Twin Cities, and other experiential learning opportunities.  Weekly seminar sessions incorporating readings, reflective writing, and individual and small group projects complement the experiential aspects of the course. As the semester progresses, each course member will develop a curricular unit aimed at teaching an important environmental issue to diverse adolescents attending urban public schools.  The curricular unit is a significant undertaking that provides students with the opportunity to synthesize all aspects of the course material in a creative, pragmatic and integrative manner.

General Education Requirements:

Distribution Requirements:
Social science

Course Materials

The structure and function of the vertebrate nervous system, with a focus on the human nervous system, will be explored through analysis of human brain specimens and sections, and artistic photographic and computer graphic representations of nervous system structures at both the microscopic and systems levels. This course is designed to provide an understanding of both the peripheral and central nervous system with a focus on brain, brainstem, and spinal cord structure and function through observation and study of normal brain tissue and discussion of clinical cases. This course is intended for students with a strong interest in neuroscience. Three lecture hours and one three-hour lab each week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

The structure and function of the vertebrate nervous system, with a focus on the human nervous system, will be explored through analysis of human brain specimens and sections, and artistic photographic and computer graphic representations of nervous system structures at both the microscopic and systems levels. This course is designed to provide an understanding of both the peripheral and central nervous system with a focus on brain, brainstem, and spinal cord structure and function through observation and study of normal brain tissue and discussion of clinical cases. This course is intended for students with a strong interest in neuroscience. Three lecture hours and one three-hour lab each week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course is an introduction to the diverse field of microbiology, including prokaryotes, archaea, viruses, and eukaryotic microorganisms. We will explore the ways microbes interact with their environments, in particular their diverse and complex relationships with humans. The course includes regular discussion of primary literature and an emphasis on written and oral communication. In the laboratory, students will practice fundamental microbiology techniques and data analysis, and lead hypotheses-driven independent investigations. Three lecture hours and one three-hour laboratory each week. Prerequisite(s): BIOL 170, BIOL 190 and BIOL 200; CHEM 211 recommended.

General Education Requirements:
Writing WA

Distribution Requirements:
Natural science and mathematics

Course Materials

This course is an introduction to the diverse field of microbiology, including prokaryotes, archaea, viruses, and eukaryotic microorganisms. We will explore the ways microbes interact with their environments, in particular their diverse and complex relationships with humans. The course includes regular discussion of primary literature and an emphasis on written and oral communication. In the laboratory, students will practice fundamental microbiology techniques and data analysis, and lead hypotheses-driven independent investigations. Three lecture hours and one three-hour laboratory each week. Prerequisite(s): BIOL 170, BIOL 190 and BIOL 200; CHEM 211 recommended.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of the cellular and molecular mechanisms underlying the biological basis of behavior. While particular emphasis is placed on the molecular and cellular components of the nervous system, these components are the foundation for the analysis of various systems. Discussion topics may include the role of neurotransmitters, neuromodulators and receptors in learning and memory, multiple sclerosis, Alzheimer's disease and drug addiction. The laboratory will be used to introduce major research techniques in neurobiology. These techniques will be used in independently designed research projects. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This interdisciplnary course will examine selected topics in computational biology, including basic bioinformatics, algorithms used in genomics and genome analysis, computational techniques for systems biology, and synthetic biology. Prerequisite(s): COMP 123; BIOL 190 recommended

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

An exploration of one of the central organizing ideas of modern biology, the theory of evolution. Topics that will be covered include natural and sexual selection, adaptation, comparative methods, phylogeny, speciation, population genetics, molecular evolution, the origin of life, and others. The course will consist of lectures and discussions based on readings drawn from a variety of sources with an emphasis on primary literature. Three lecture hours each week. Prerequisite(s): BIOL 180 and BIOL 190

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

A study of biological processes at the molecular level with an emphasis on the chemistry of biological molecules, elements of physical biochemistry, the structure of proteins, the mechanisms and kinetics of enzyme catalyzed reactions, and selected topics in intermediary metabolism, including the metabolism of carbohydrates and lipids. Three lecture hours and one three-hour laboratory per week. Prerequisite(s): BIOL 200 and CHEM 212, or permission of the instructor. Students must earn a C- or higher in prerequisite courses.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course is an introduction to vertebrate immunity. Its evolution, cellular and molecular mechanisms, health and disease functions and therapeutic manipulations are explored through approaches including lectures, clinical case studies, extensive reading of the primary literature, problem-solving and an intensive focus on scientific writing. The course typically includes a civic engagement component where students work with health organizations and schools in the Twin Cities area. The course laboratory uses guided exercises and independent projects to focus on current immunological techniques including flow cytometry, magnetic cell sorting and antibody assays. Three hours of lectures and four to six hours of laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200; CHEM 112; junior or senior standing

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course is an introduction to vertebrate immunity. Its evolution, cellular and molecular mechanisms, health and disease functions and therapeutic manipulations are explored through approaches including lectures, clinical case studies, extensive reading of the primary literature, problem-solving and an intensive focus on scientific writing. The course typically includes a civic engagement component where students work with health organizations and schools in the Twin Cities area. The course laboratory uses guided exercises and independent projects to focus on current immunological techniques including flow cytometry, magnetic cell sorting and antibody assays. Three hours of lectures and four to six hours of laboratory per week. Prerequisite(s): BIOL 190 and BIOL 200; CHEM 112; junior or senior standing

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

The Arctic is Earth's most rapidly warming region. It is also home to massive carbon reservoirs and diverse biological adaptations to extreme elements, as well as home to Indigenous populations and the site of oil extraction and vanishing sea ice. We will examine how climate change is impacting the biodiversity, ecophysiology, and biogeochemistry of this crucial biome, and as a result, the rest of the world. As an upper-level biology course, Arctic Ecology aims to challenge students to improve their science communication skills through varied written, spoken, and visual presentations. Students will also be challenged to synthesize content across systems and create novel hypotheses about current and future impacts of change at a species, community, ecosystem, and landscape scale. Three lecture hours each week. Prerequisite(s): BIOL 170

General Education Requirements:
Writing WP

Distribution Requirements:
Natural science and mathematics

Course Materials

This course aims to integrate organismal, cellular, genetic and molecular approaches to the study of animal development. We will analyze a diversity of mechanisms, ranging from ones that set up pattern formation in the unfertilized egg to those governing morphogenesis of organ systems. Evolution of developmental mechanisms will also be discussed. The lab component will incorporate both descriptive and experimental embryological techniques. Three hours of lecture and one three-hour laboratory each week. Prerequisite(s): BIOL 180, BIOL 190 and BIOL 200, or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

This course aims to integrate organismal, cellular, genetic and molecular approaches to the study of animal development. We will analyze a diversity of mechanisms, ranging from ones that set up pattern formation in the unfertilized egg to those governing morphogenesis of organ systems. Evolution of developmental mechanisms will also be discussed. The lab component will incorporate both descriptive and experimental embryological techniques. Three hours of lecture and one three-hour laboratory each week. Prerequisite(s): BIOL 180, BIOL 190 and BIOL 200, or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Why do animals behave the way they do? Why do lions have manes while leopards don’t? Why do elephants and bees live in groups but many other species do not? Why does your friendly neighborhood squirrel get so busy late in the fall and again in the spring? Why do certain wolves ‘fish’ but others never learn the technique? In this course we will explore the fundamentals of animal behavior and use that foundation to understand how we can better manage and conserve animal biodiversity. Labs will include hands-on experience with tools that will allow us to quantify behavior, develop ethograms, and understand species’ repertoires. Three hours of lecture/discussion and three hours of laboratory each week. Prerequisites: BIOL/ENVI 170 and either BIOL180 or ENVI 240. Recommended: STAT 155.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Our brains are tightly linked to the everyday inner-workings of our bodies. This course will focus on the field of neuroendocrinology: how the nervous system communicates with distal tissues of the body via neuropeptides and hormones. Topics include stress, social and sexual behaviors, growth and development, homeostatic regulation, hunger and feeding, and sleep. Class periods will involve interactive lectures and discussions of primary neuroendocrinology literature. Laboratory periods will apply various behavioral and molecular techniques to student-designed projects. Prerequisites: BIOL 190 and 200.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

Varies by semester. Consult the department or class schedule for current listing.

General Education Requirements:

Distribution Requirements:

Course Materials

Varies by semester. Consult the department or class schedule for current listing.

General Education Requirements:

Distribution Requirements:

Course Materials

This seminar course centers on our understanding of the molecular events underlying the development of various cancers, including cell cycle control, DNA damage, tumor microenvironment, as well as therapeutic approaches to treatment, and associated difficulties. Two major goals of the course include developing an ability to understand cancerous transformation and learning to critically evaluate literature in the field. We will discuss primary articles and secondary reviews in a roundtable format, and students will present written and oral critiques of the literature and develop a final individual project. Three lecture/discussion hours each week. Prerequisite(s): BIOL 200, and junior or senior standing; or permission of instructor.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

As we usher into the Anthropocene, where ‘Amazon’ is recognized as a global marketing forum more than the mighty river, the fate of biodiversity reflects a similar trajectory. Human population growth has resulted in unprecedented changes in our planet’s ecosystems. Species have been lost, modified, and forced to live in concrete jungles. Human history, culture and socio-political nuances in different parts of the world result in distinctive challenges as well as enigmatically optimistic scenarios with respect to wildlife conservation. In this seminar we will investigate perspectives from South Asia where animals and humans coexist at very high density and proximity, consider the North American system where views about biodiversity can often be very polarizing, and explore conservation in Africa where pockets of pristine wilderness persist. We will engage with the primary literature of wildlife conservation, hear from scientists on the frontlines, and discuss the idea that there could be a unifying model that safeguards biodiversity without detrimentally impeding human development. Three hours of lecture/discussion each week. Prerequisites: BIOL/ENVI 170 and either BIOL 180 or ENVI 240.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials

What exactly happens inside the brain when the mind goes awry? What are the subjective experiences of psychiatric disorders and how can our understanding of the brain inform treatments for these conditions? Today, modern neuroscience has greatly advanced our insight into these questions. This course is designed to overview major types of mental illness and delve into their respective neurobiological origins. To explore these topics, class will involve (1) reflecting on first-hand narratives of individuals with mental illness, (2) engaging in interactive lectures, during which you’ll acquire and share knowledge, and (3) contributing to discussions on neuroscience research articles. A multi-draft paper on a topic of your choosing is optional.

General Education Requirements:

Distribution Requirements:
Natural science and mathematics

Course Materials